Dr Emily Bethell

Journal article

Howarth E, Witham C, Bethell E. 2025. Does Measuring Social Attention Lead to Changes in Behavior? A Preliminary Investigation into the Implications of Attention Bias Trials on Behavior in Rhesus Macaques Journal of Applied Animal Welfare Science, :1-14 DOI Author Url Publisher Url Public Url

Arbuckle K, Bethell EJ, Hawthorn DJ, Hunt K, Khera M, Lewis Z, Mitchell J, Nicholl MH, Reynolds LA. 2025. Low socioeconomic status is an under-recognised source of challenges in academia Journal of Zoology, DOI Publisher Url Public Url

Howarth ERI, Szott ID, Witham CL, Wilding CS, Bethell EJ. 2023. Genetic polymorphisms in the serotonin, dopamine and opioid pathways influence social attention in rhesus macaques (Macaca mulatta) Singh AS. PLoS One, 18 :1-29 DOI Author Url Publisher Url Public Url

Wilson VAD, Bethell EJ, Nawroth C. 2023. The use of gaze to study cognition: limitations, solutions, and applications to animal welfare Frontiers in Psychology, 14 :1-10 DOI Author Url Publisher Url Public Url

Bethell EJ, Khan W, Hussain A. 2022. A deep transfer learning model for head pose estimation in rhesus macaques during cognitive tasks: towards a nonrestraint noninvasive 3Rs approach Applied Animal Behaviour Science, DOI Publisher Url Public Url

Cassidy LC, Bethell EJ, Brockhausen RR, Boretius S, Treue S, Pfefferle D. 2021. The dot-probe attention bias task as a method to assess psychological wellbeing after anesthesia: A study with adult female long-tailed macaques (Macaca fascicularis). European Surgical Research, DOI Author Url Publisher Url Public Url

Evans R, Lyons M, Brewer G, Bethell E. 2021. A domestic cat (Felis silvestris catus) model of triarchic psychopathy factors: Development and initial validation of the CAT-Tri+ questionnaire Journal of Research in Personality, 95 :1-10 DOI Author Url Publisher Url Public Url

Eccles GR, Bethell EJ, Greggor AL, Mettke-Hofmann C. 2021. Individual Variation in Dietary Wariness Is Predicted by Head Color in a Specialist Feeder, the Gouldian Finch Frontiers in Ecology and Evolution, 9 DOI Author Url Publisher Url Public Url

Crump A, Bethell E, Arnott G. 2021. Optimism and pasture access in dairy cows Scientific Reports, 11 DOI Author Url Publisher Url Public Url

Bethell EJ. 2021. Studying Primates: How to Design, Conduct and Report Primatological Research - JM Setchell (2019). Published by Cambridge University Press, Shaftesbury Road, Cambridge CB2 8BS, UK. 362 pages Paperback (ISBN: 978-1108434270). Price £26.99. Animal Welfare, 30 :106-108 DOI Publisher Url

Howarth ER, Kemp C, Thatcher HR, Szott ID, Farningham D, Witham CL, Holmes A, Semple S, Bethell EJ. 2021. Developing and validating attention bias tools for assessing trait and state affect in animals: A worked example with Macaca mulatta Applied Animal Behaviour Science, 234 DOI Author Url Publisher Url Public Url

Crump A, Bethell EJ, Earley R, Lee VE, Mendl M, Oldham L, Turner SP, Arnott G. 2020. Emotion in animal contests Proceedings of the Royal Society B: Biological Sciences, 287 :20201715-20201715 DOI Author Url Publisher Url Public Url

Crump A, Bethell EJ, Earley R, Lee VE, Mendl M, Oldham L, Turner SP, Arnott G. 2020. Emotion in animal contests Proceedings of the Royal Society B: Biological Sciences, 287 DOI

Mettke-Hofmann C, Eccles G, Greggor A, Bethell EJ. 2020. Cognition in a changing world: Red-headed Gouldian finches enter spatially unfamiliar habitats more readily than do black-headed birds Frontiers in Ecology and Evolution, 8 DOI Author Url Publisher Url Public Url

Crump A, Mullens C, Bethell EJ, Cunningham EM, Arnott G. 2020. Microplastics disrupt hermit crab shell selection Biology Letters, 16 DOI Author Url Publisher Url Public Url

Bethell EJ. 2019. Pasture Access Affects Behavioral Indicators of Wellbeing in Dairy Cows Animals, 9 DOI Author Url Publisher Url Public Url

Bethell E, Cassidy LC, Brockhausen RR, Pfefferle D. 2019. Toward a Standardised Test of Fearful Temperament in Primates: a Sensitive Alternative to the Human Intruder Task for Laboratory-Housed Rhesus Macaques (Macaca mulatta) Frontiers in Psychology, 10 DOI Author Url Publisher Url Public Url

Bethell E, Kemp C, Thatcher H, Schroeder J, Arbuckle K, Farningham D, Witham C, Holmes A, MacLarnon A, Semple S. 2019. Heritability and maternal effects on social attention during an attention bias task in a non-human primate, Macaca mulatta EcoEvoRxiv Preprints, DOI Publisher Url

Crump A, Arnott G, Bethell EJ. 2018. Affect-Driven Attention Biases as Animal Welfare Indicators: Review and Methods Animals, 8 DOI Author Url Publisher Url Public Url

Cronin KA, Bethell EJ, Jacobson SL, Egelkamp C, Hopper LM, Ross SR. 2018. Evaluating Mood Changes in Response to Anthropogenic Noise with a Response-Slowing Task in Three Species of Zoo-housed Primates Animal Behavior & Cognition, 5 :209-221 DOI Publisher Url Public Url

Kemp C, Thatcher H, Farningham D, Witham C, MacLarnon A, Holmes A, Semple S, Bethell EJ. 2017. A protocol for training group-housed rhesus macaques (Macaca mulatta) to cooperate with husbandry and research procedures using positive reinforcement Applied Animal Behaviour Science, 197 :90-100 DOI Author Url Publisher Url Public Url

Lyons MT, Brewer G, Bethell EJ. 2016. Sex-Specific Effect of Recalled Parenting on Affective and Cognitive Empathy in Adulthood Current Psychology, DOI Author Url Publisher Url Public Url

Bethell EJ, Holmes A, MacLarnon A, Semple S. 2016. Emotion evaluation and response slowing in a non-human primate: New directions for cognitive bias measures of animal welfare Behavioral Sciences, 6 DOI Author Url Publisher Url Public Url

Bethell EJ. 2015. A “How-To” Guide for Designing Judgment Bias Studies to Assess Captive Animal Welfare Journal of Applied Animal Welfare Science, 18 :S18-S42 DOI Author Url Publisher Url Public Url

Bethell EJ, Koyama NF. 2015. Happy hamsters? Enrichment induces positive judgement bias for mildly (but not truly) ambiguous cues to reward and punishment in Mesocricetus auratus. Royal Society Open Science, 2 :1-17 DOI Author Url Publisher Url Public Url

Jonason PK, Lyons M, Bethell EJ. 2014. The making of Darth Vader: Parent–child care and the Dark Triad Personality and Individual Differences, 67 :30-34 DOI Author Url Publisher Url

Jonason PK, Lyons M, Bethell EJ, Ross R. 2013. Different routes to limited empathy in the sexes: Examining the links between the Dark Triad and empathy Personality and Individual Differences, 54 :572-576 DOI

Bethell EJ, Holmes A, Maclarnon A, Semple S. 2012. Evidence that emotion mediates social attention in rhesus macaques. PLoS One, 7 :e44387-e44387 DOI Author Url Publisher Url Public Url

Bethell EJ, Holmes A, MacLarnon A, Semple S. 2012. Cognitive bias in a non-human primate: Husbandry procedures influence cognitive indicators of psychological well-being in captive rhesus macaques Animal Welfare, 21 :185-195 DOI Author Url Publisher Url Public Url

Bethell EJ, Vick S-J, Bard KA. 2007. Measurement of eye-gaze in chimpanzees (Pan troglodytes) American Journal of Primatology, 69 :562-575 DOI Author Url Publisher Url

Bethell EJ, Semple S, MacLarnon A, Holmes A. 2007. The effect of emotion state on responses to social stimuli by rhesus macaques Primate Eye, 92 :5-6

Bethell EJ, Waller BM. 2005. Altruism or cooperation in captive chimpanzees, Pan troglodytes? Folia Primatologica, 76 :242-244 DOI Author Url Publisher Url

Bethell EJ, Whiten A, Muhumuza G, Kakura J. 2000. Active plant food division and sharing by wild chimpanzees Primate Report, 56 :67-71

Crump A, Jenkins K, Bethell EJ, Ferris CP, Arnott G. Pasture access and eye temperature in dairy cows Journal of Applied Animal Welfare Science, DOI Author Url Publisher Url Public Url

Chapters

Bethell EJ, Pfefferle D. 2023. Cognitive bias tasks: A new set of approaches to assess welfare in nonhuman primates Robinson LM, Weiss A. Nonhuman Primate Welfare: From History, Science, and Ethics to Practice :207-230 Springer International Publishing 9783030827076 DOI Publisher Url Public Url

Bethell EJ. 2021. Color Red Encyclopedia of Evolutionary Psychological Science :1187-1189 Springer International Publishing 9783319196497 DOI Publisher Url

Editorial/letter to the editor

Wilson VAD, Bethell EJ, Nawroth C. 2023. Editorial: Using gaze to study social knowledge: current challenges and future directions Frontiers in Psychology, 14 DOI Author Url Publisher Url

Bethell EJ, Dean L, Smith A, Bearder SK. 2018. Celebrating 50 years of the Primate Society of Great Britain Evolutionary Anthropology, 27 :74-77 DOI Publisher Url

Bethell EJ. 2016. Color Red Shackelford TK, Weekes-Shackelfrod VA. :1-3 DOI

Book review

Bethell EJ. 2021. Studying Primates: How to Design, Conduct and Report Primatological Research ANIMAL WELFARE, 30 :106-108 Author Url

Bethell E. 2017. Book review: Being Jane Primate Eye,

Dataset

Witham C, Bethell E. 2019. The Macaque Stimulus Set DOI Publisher Url

Internet publication

Prescott M, Bethell EJ, Kemp C. 2015. NC3Rs Macaque website Author Url Publisher Url

Books (authored)

Bethell E, Milsom C. 2014. Posters and Presentations :1-144 Palgrave Macmillan. London 9781137357083 DOI Publisher Url

Williams K, Bethell EJ, Lawton J, Parfitt-Brown C, Richardson M, Rowe V. 2011. Completing your PhD Palgrave MacMillan. London 978-0230292819 DOI Publisher Url

Williams K, Bethell E, Lawton J, Parfitt C, Richardson M, Rowe V. 2010. Planning Your PhD Palgrave MacMillan 9780230251939 DOI Publisher Url

Conference publication

Kemp C, Bethell EJ. Measuring psychological well-being in captive rhesus macaques: attention bias http://behaviour-2015.m.asnevents.com.au/schedule/session/6854/abstract/24936, Behaviour

Professional activities

Research Grants Awarded:

Duration of research project: 4 years, Grant value (£): 575000, European Cooperation in Science and Technology, Tom Smulders, University of Newcastle, CA23106 - An Evolutionary View to Understanding Affective States across Species (AFFECT-EVO). 2024

Media Coverage:

NY TImes Comment on conrtolling macaques during G20 summit. 2023
www.proquest.com
www.nytimes.com

Comment in The Guardian on research in animal cognition and welfare 2021

News outlet coverage of paper - International (India, Russia, Ireland) 2020
www.indiatimes.com
www.popmech.ru
www.independent.ie

Media coverage in 114 media news outlets for Microplastics disrupt hermit crab shell selection 2020
www.independent.co.uk
eandt.theiet.org
www.theguardian.com

Daily Mail

Discovery Magazine

Media coverage of Royal Soc paper 2015 in Science, Discovery, Daily Mail
www.dailymail.co.uk
news.sciencemag.org

Media coverage of Royal Society paper 2015: Metro, USA Today, Huffington Post
www.usatoday.com
www.huffingtonpost.com
metro.co.uk

Media/radio coverage of Royal Society paper 2015 the Times, BBC Radio, Liverpool Echo
naked_scientists_popup_player.html
www.liverpoolecho.co.uk
www.thetimes.co.uk

Conference presentation:

Oral presentation, LNG Christmas Seminar, The gut-brain axis in animal welfare. 2021

Oral presentation, DPZ, Gottingen, Germany, Gottingen University Seminar Series, Invited Seminar Speaker. 2018

Oral presentation, Queens Belfast, Queens Belfast Seminar, Invited Seminar Speaker. 2018

Oral presentation, Imperial College. Silwood Campus, London, Imperial College Seminar, Invited Seminar Speaker. 2017

Oral presentation, India, Venus International, Invited Keynote. 2017

Oral presentation, Japan, International congress for Psychology, Invited: Measuring animal emotions. 2016

Oral presentation, LJMU, LJMU Postgraduate Research Day, Invited speaker: 3Rs in primate research. 2016

Oral presentation, Liverpool, LivLABS, Introduction to the Liverpool League of Animal Behaviour Scientists. 2016

Oral presentation, Rome, Italy, European Federation for Primatology, Cognitive bias - methods for measuring emotion in zoo animals. 2015

Oral presentation, Australia, Behaviour, Measuring psychological well-being in captive rhesus macaques: attention bias. 2015

Poster presentation, Roehampton, London, Primate Society of Great Britain spring conference, Effects of a habituation regime on attention bias for veterinary stimuli in macaques. 2015

Poster presentation, Roehampton, London, Primate Society of Great Britain spring conference, What is neutral? Attention bias for facial expressions in macaques. 2015

Oral presentation, Newcastle, Experimental Psychology Society, Invited: Attention bias a cross species comparison. 2014

Oral presentation, London, NC3RS Primate Welfare workshop, Attention bias: developing a methods to asses wellbeing in macaque breeding facilities. 2014

Conference organisation:

Organising Committee, European Conference for Behavioural Biology, https://www.ljmu.ac.uk/conferences/ecbb. 2018

PSGB Meetings Organiser, PSGB Spring meeting 2018. 2018

PSGB Meetings Organiser, Primate Society of Great Britain Spring Conference 2017. 2017

Scientific Meetings Coordinator for the PSGB, Primate Society of Great Britain spring conference 2016, http://www.psgb.org/. 2016

Chair of Organising Committee, Primate Society of Great Britain 50th anniversary conference. 2015

Award:

Venus International, Outstanding Scientist Award for contributions to field of primate behaviour and cognition, https://venusinfo.org/research/arm/2017/info.html. 2017

LJMU Student Union, Best Teaching Team. 2011

Association for the Study of Animal Behaviour, Honourable mention in student competition for best talk. 2007

Primate Society of Great Britain, Best Student Presentation. 2007

Membership of professional bodies:

Animal Welfare Research Network, Member. 2016

Universities Federation of Animal Welfare, Member. 2004

Association for the Study of Animal Behaviour, Member. 2000

Association for Psychological Science, Member.

Primate Society of Great Britain, Member.

Editorial boards:

Review Editor in Comparative Psychology, Frontiers in Psychology, http://journal.frontiersin.org/journal/psychology#editorial-board. 2015

External committees:

Primate Society of Great Britain, Chair of Organising Committee, 50th anniversary conference organising committee. 2015

PSGB, PSGB Scientific Meetings Coordinator. 2015

Primate Society of Great Britain, member, Captive Care Working Party, http://www.psgb.org/. 2015

Other invited event:

Rome, Invited speaker, Invited symposium speaker: European Federation of Primatology 2015. 2015

Stirlin University, Invited speaker, Psychology Research Seminar. 2015

Detroit Zoological Society, Detroit, USA, Invited speaker, Zoo Animal Welfare Symposium. 2014

Newcastle University, Invited speaker, Experimental Psychology Society. 2014

University of Stirling, Invited seminar speaker - psychology Department, Stirling, University of Stirling Psychology Seminar Series. 2012

Birkbeck College, London, Invited paper, Birkbeck College Seminar Series. 2010

London, UK, Invited paper presented, NC3Rs Primate Welfare meeting. 2010

UCL, London, Invited paper presented, University College London Anthropology seminar series. 2010

Fellowships:

Liverpool John Moores University, LJMU ECR Fellowship. 2013

Advance HE, Fellow of the HEA. 2012